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Northampton’s Tommy Freeman takes Jack Nowell’s place on the wing in one of four changes for England’s match against South Africa on Saturday.
Freeman, who impressed on the summer tour of Australia, only recently made his club comeback from a foot injury.
Elsewhere Ellis Genge and Luke Cowan-Dickie have dropped to the bench with Mako Vunipola and hooker Jamie George coming into the starting line-up.
Alex Coles replaces Sam Simmonds at flanker to add a third line-out jumper.
Jack van Poortvliet continues at scrum-half ahead of Leicester team-mate Ben Youngs.
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